(Getty) Victoria was played by Rachelle Lefevre in Twilight and New Moon. As Twilight aficionados already know, Summit Entertainment announced last night that fellow ginger gal Bryce Dallas Howard will replace Lefevre as Victoria in … Rachelle Lefevre (l.), who played Victoria in the first two 'Twilight' films, and her replacement, Bryce Dallas Howard. As for her work on the big screen, she appeared in Confessions of a Dangerous Mind as Tuvia, among other projects. This is a list of the cast members from The Twilight Saga film series, which is based on the … Prior to her role in the Twilight Saga, Lefevre had worked in various TV shows, though mostly in minor roles.